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Guanine: One of the four bases in DNA that make up the letters ATGC. Guanine is the "G". The others are adenine , cytosine , and thymine . Guanine always pairs with cytosine to form the base pair G-C (guanine-cytosine) in the DNA. The other base pair in the DNA is A-T (adenine-thymine). Each base pair forms a rung in the DNA ladder. Guanine is also one of the four bases in RNA .

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